This work was supported by the Korea Health Technology R&D Project, funded by the Ministry of Health & Welfare, Republic of Korea (HI15C2817), Ministry of Health & Welfare (government – wide R&D fund project for infectious disease research (HG18C0029)) to Y.K.C. 
B.B is supported by a Wellcome Trust/Royal Society Sir Henry Dale Fellowship (210462/Z/18/Z) and previously a Wellcome Trust Senior Investigator Award (099220/Z/12/Z) to R.M. Elliott. 
J.U.J is supported by awards from the National Institute of Health (AI116585, AI129496, AI40718 and AI140705).